In the world of fungal civilization, where forests of giant mushrooms cover the land and messages are transmitted through mycelial networks, gravity is closely linked to personal confidence. The cloud shepherd Zella was murdered by his own shadow after working at a high altitude. She discovers that Shadow has awakened an independent consciousness and escaped into the underground mycelium labyrinth, and her memories and emotions are being lost with it. In order to retrieve the "anchor" of her soul, Zella fell from the sky into the bottomless world of fungal roots, only to discover that her shadow's rebellion was just the prelude to a huge conspiracy - the mycelial network matrix on which the entire civilization depends for survival is secretly planning to devour all human shadows to complete its ultimate evolution. Zeira must form an alliance with the dark half who betrayed her, and fight to save all of her "whole" selves under the strange laws of faith and gravity.